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A new species of Dyrosaurus (Crocodylomorpha, Dyrosauridae) from the early Eocene of Morocco: phylogenetic implications

Identifieur interne : 007035 ( Main/Exploration ); précédent : 007034; suivant : 007036

A new species of Dyrosaurus (Crocodylomorpha, Dyrosauridae) from the early Eocene of Morocco: phylogenetic implications

Auteurs : Stéphane Jouve [France] ; Mohamed Iarochène [Maroc] ; Baâdi Bouya [Maroc] ; Mbarek Amaghzaz [Maroc]

Source :

RBID : ISTEX:A9014BF6D0BE876CF0D2360E568EBB1A1E93C436

Descripteurs français

English descriptors

Abstract

A new dyrosaurid is described from the Ypresian of the phosphatic deposits of the Oulad Abdoun Basin of Morocco. It is based on numerous cranial and postcranial remains, allowing an almost complete reconstruction. This new Dyrosaurus species, Dyrosaurus maghribensis sp. nov., is currently only known from Morocco. It differs from D. phosphaticus, present in contemporaneous levels of Algeria and Tunisia, by several autapomorpies, including a smooth dorsal margin of the parietal and widely opened choanae. A phylogenetic analysis, using 47 taxa and 234 morphological characters, shows the dyrosaurids as the sister taxon of pholidosaurids, which include Elosuchus, Sarcosuchus, Terminonaris and Pholidosaurus, and the thalattosuchians. Goniopholididae is a non‐monophyletic group; however, if dyrosaurids are not included in the analysis, the result differs and the goniopholidids form a distinct clade. If Thalattosuchia is excluded, both Goniopholididae and Pholidosauridae become paraphyletic assemblages. Thus, phylogenetic problems remain with respect to longirostrine clade, and more attention should be paid to resolving their evolutionary relationships amongst the crocodyliforms. © 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 148, 603–656.

Url:
DOI: 10.1111/j.1096-3642.2006.00241.x


Affiliations:


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Le document en format XML

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<keywords scheme="KwdEn" xml:lang="en">
<term>Abdoun</term>
<term>Abhandlungen</term>
<term>Alveolus</term>
<term>American museum novitates</term>
<term>Anatosuchus</term>
<term>Anterior</term>
<term>Anterior contour</term>
<term>Anterior edge</term>
<term>Anterior margin</term>
<term>Anterior process</term>
<term>Anterior view</term>
<term>Anteriorly</term>
<term>Anteriormost</term>
<term>Anterodorsal</term>
<term>Anterolateral</term>
<term>Anterolateral postorbital process</term>
<term>Anteromedial</term>
<term>Anteromedial margin</term>
<term>Anteroposterior</term>
<term>Anteroposteriorly</term>
<term>Anteroventral</term>
<term>Anteroventrally</term>
<term>Antorbital</term>
<term>Araripesuchus</term>
<term>Archosauria</term>
<term>Arech</term>
<term>Articular</term>
<term>Articular glenoid cavity</term>
<term>Articular surface</term>
<term>Attened</term>
<term>Basal</term>
<term>Basioccipital</term>
<term>Basioccipital tubera</term>
<term>Basisphenoid</term>
<term>Basisphenoid rostrum</term>
<term>Benton</term>
<term>Benton clark</term>
<term>Bequaerti</term>
<term>Bernissartia</term>
<term>Brinkman</term>
<term>Brochu</term>
<term>Broin</term>
<term>Buckley brochu</term>
<term>Buffetaut</term>
<term>Buscalioni</term>
<term>Calsoyasuchus</term>
<term>Calvo</term>
<term>Caudal</term>
<term>Centrum</term>
<term>Cervical</term>
<term>Cervicals</term>
<term>Chenanisuchus</term>
<term>Chenanisuchus lateroculi</term>
<term>Chimaerasuchus</term>
<term>Choanae</term>
<term>Clade</term>
<term>Comahuesuchus</term>
<term>Comptes</term>
<term>Comptes rendus</term>
<term>Concavity</term>
<term>Condyle</term>
<term>Consensus tree</term>
<term>Coracoid</term>
<term>Coronoid</term>
<term>Cranial</term>
<term>Cretaceous</term>
<term>Crocodile</term>
<term>Crocodilia</term>
<term>Crocodilian</term>
<term>Crocodiliens</term>
<term>Crocodyliform</term>
<term>Crocodyliformes</term>
<term>Crocodyliforms</term>
<term>Crocodylomorph</term>
<term>Crocodylomorpha</term>
<term>Crocodylus</term>
<term>Dentary</term>
<term>Dentary teeth</term>
<term>Diapophysis</term>
<term>Dibothrosuchus</term>
<term>Dibothrosuchus protosuchus orthosuchus gobiosuchus notosuchus sebecus uruguaysuchus</term>
<term>Dollo</term>
<term>Dorsal</term>
<term>Dorsal armour</term>
<term>Dorsal margin</term>
<term>Dorsal osteoderms</term>
<term>Dorsal part</term>
<term>Dorsal surface</term>
<term>Dorsal vertebrae</term>
<term>Dorsal view</term>
<term>Dorsally</term>
<term>Dorsals</term>
<term>Dorsoventrally</term>
<term>Dyrosaurid</term>
<term>Dyrosauridae</term>
<term>Dyrosaurids</term>
<term>Dyrosaurus</term>
<term>Dyrosaurus maghribensis</term>
<term>Dyrosaurus phosphaticus</term>
<term>Ectopterygoid</term>
<term>Elosuchus</term>
<term>Eocene</term>
<term>Eustachian</term>
<term>Eutretauranosuchus</term>
<term>Exoccipital</term>
<term>External mandibular fenestra</term>
<term>External nares</term>
<term>External naris</term>
<term>Femur</term>
<term>Fenestra</term>
<term>Foramen</term>
<term>Foramen magnum</term>
<term>Fossa</term>
<term>Gado</term>
<term>Gasparini</term>
<term>Gavialis</term>
<term>Gavialis gangeticus</term>
<term>Glenoid</term>
<term>Glenoid fossa</term>
<term>Gobiosuchus</term>
<term>Gomani</term>
<term>Goniopholis</term>
<term>Goniopholis simus</term>
<term>Halstead</term>
<term>Halstead middleton</term>
<term>Holotype</term>
<term>Hsisosuchus</term>
<term>Humerus</term>
<term>Hyleochampsa</term>
<term>Hypapophyses</term>
<term>Hypapophysis</term>
<term>Hyposaurus</term>
<term>Ianwilsoni</term>
<term>Ilium</term>
<term>Imperator</term>
<term>Infratemporal</term>
<term>Infratemporal fenestra</term>
<term>Infratemporal fenestrae</term>
<term>Interfenestral</term>
<term>Internal nares</term>
<term>Ischium</term>
<term>Jouve</term>
<term>Jouve schwarz</term>
<term>Jugal</term>
<term>Jurassic</term>
<term>Lacrimal</term>
<term>Lamina</term>
<term>Larsson</term>
<term>Larsson gado</term>
<term>Late cretaceous</term>
<term>Lateral</term>
<term>Lateral eustachian foramen</term>
<term>Lateral margin</term>
<term>Lateral margins</term>
<term>Lateral surface</term>
<term>Lateral view</term>
<term>Laterally</term>
<term>Lateroculi</term>
<term>Laterosphenoid</term>
<term>Libycosuchus</term>
<term>Linnean</term>
<term>Linnean society</term>
<term>Longirostrine</term>
<term>Longirostrine forms</term>
<term>Longirostrine morphology</term>
<term>Longitudinal rows</term>
<term>Lower cretaceous</term>
<term>Lower eocene</term>
<term>Lpup lppm</term>
<term>Maghribensis</term>
<term>Malawisuchus</term>
<term>Mandible</term>
<term>Mandibular</term>
<term>Mandibular symphysis</term>
<term>Margin</term>
<term>Maroc</term>
<term>Maxilla</term>
<term>Maxillary</term>
<term>Maxillary teeth</term>
<term>Maxillary tooth</term>
<term>Medial</term>
<term>Medial eustachian foramen</term>
<term>Medial view</term>
<term>Medially</term>
<term>Mera</term>
<term>Mnhn</term>
<term>Monophyly</term>
<term>Mook</term>
<term>Naris</term>
<term>Nasal</term>
<term>Natural history</term>
<term>Neosuchians</term>
<term>Neural</term>
<term>Neural arch</term>
<term>Neural arches</term>
<term>Neural spine</term>
<term>Neural spines</term>
<term>Notosuchia</term>
<term>Notosuchus</term>
<term>Observ</term>
<term>Occipital</term>
<term>Occipital condyle</term>
<term>Occipital face</term>
<term>Occipital tuberosities</term>
<term>Occipital tuberosity</term>
<term>Occipital view</term>
<term>Odontoid process</term>
<term>Orientated</term>
<term>Ortega</term>
<term>Orthosuchus</term>
<term>Osteoderm</term>
<term>Osteoderms</term>
<term>Other crocodyliformes</term>
<term>Other crocodyliforms</term>
<term>Other dyrosaurids</term>
<term>Other longirostrine forms</term>
<term>Otoccipital</term>
<term>Oulad</term>
<term>Oulad abdoun basin</term>
<term>Palaeontology</term>
<term>Palatal</term>
<term>Palatine</term>
<term>Palatine fenestra</term>
<term>Paleontology</term>
<term>Parietal</term>
<term>Paroccipital</term>
<term>Paroccipital process</term>
<term>Patagonicus lomasuchus theriosuchus goniopholis bernissartia hyleochampsa terminonaris pholidosaurus sarcosuchus elosuchus pelagosaurus vectisuchus steneosaurus libycosuchus sunosuchus metriorhynchus dyrosaurus gavialis crocodylus sphenosuchus stolokrosuchus hsisosuchus shamosuchus sichuanosuchus shantungosuchus simosuchus anatosuchus calsoyasuchus rugosuchus alligator kayentasuchus comahuesuchus malawisuchus peirosaurus bretesuchus eutretauranosuchus chenanisuchus sphagesaurus chimaerasuchus</term>
<term>Pelagosaurus</term>
<term>Pers</term>
<term>Pholidosaurus</term>
<term>Phosphaticus</term>
<term>Phylogenetic</term>
<term>Phylogenetic analyses</term>
<term>Phylogenetic analysis</term>
<term>Phylogenetic implications</term>
<term>Phylogenetic position</term>
<term>Phylogenetic relationships</term>
<term>Postcranial</term>
<term>Posterior</term>
<term>Posterior edge</term>
<term>Posterior margin</term>
<term>Posterior margins</term>
<term>Posterior process</term>
<term>Posterior wall</term>
<term>Posteriorly</term>
<term>Posterodorsal</term>
<term>Posterodorsal margin</term>
<term>Posterodorsally</term>
<term>Posterolateral</term>
<term>Posteromedial</term>
<term>Posteromedial process</term>
<term>Posteroventral</term>
<term>Posteroventrally</term>
<term>Postorbital</term>
<term>Postorbital process</term>
<term>Prefrontal</term>
<term>Prefrontal pillar</term>
<term>Prefrontal pillars</term>
<term>Premaxilla</term>
<term>Premaxillary</term>
<term>Present work</term>
<term>Previous investigations</term>
<term>Protosuchus</term>
<term>Proximal head</term>
<term>Pterygoid</term>
<term>Pterygoids</term>
<term>Pubis</term>
<term>Quadrate</term>
<term>Quadrate condyle</term>
<term>Quadratojugal</term>
<term>Rendus</term>
<term>Reptile</term>
<term>Retroarticular</term>
<term>Retroarticular process</term>
<term>Rhabdognathus</term>
<term>Robust</term>
<term>Rogersii</term>
<term>Rostrum</term>
<term>Sacral</term>
<term>Same level</term>
<term>Sanz</term>
<term>Sarcosuchus</term>
<term>Sarcosuchus imperator</term>
<term>Scapula</term>
<term>Schwarz</term>
<term>Sciences naturelles</term>
<term>Sebecus</term>
<term>Sereno</term>
<term>Shantungosuchus</term>
<term>Sichuanosuchus</term>
<term>Simosuchus</term>
<term>Skull</term>
<term>Skull length</term>
<term>Skull roof</term>
<term>Snout</term>
<term>Sphagesaurus</term>
<term>Sphenosuchus</term>
<term>Squamosal</term>
<term>Steneosaurus</term>
<term>Stolokrosuchus</term>
<term>Storrs</term>
<term>Suborbital</term>
<term>Suborbital fenestra</term>
<term>Sunosuchus</term>
<term>Sunosuchus junggarensis</term>
<term>Supraoccipital</term>
<term>Supratemporal</term>
<term>Supratemporal fenestra</term>
<term>Supratemporal fenestrae</term>
<term>Surangular</term>
<term>Suture</term>
<term>Sutured</term>
<term>Symphysis</term>
<term>Taquet</term>
<term>Taxon</term>
<term>Temporal canal</term>
<term>Terminonaris</term>
<term>Thalattosuchia</term>
<term>Thalattosuchians</term>
<term>Theriosuchus</term>
<term>Transverse process</term>
<term>Transverse section</term>
<term>Troxell</term>
<term>Tubera</term>
<term>Tuberosity</term>
<term>Turner calvo</term>
<term>Tykoski</term>
<term>Unpublished dphil thesis</term>
<term>Upper cretaceous</term>
<term>Upper triassic</term>
<term>Uruguaysuchus</term>
<term>Vectisuchus</term>
<term>Vectisuchus leptognathus</term>
<term>Ventral</term>
<term>Ventral edge</term>
<term>Ventral margin</term>
<term>Ventral part</term>
<term>Ventral portion</term>
<term>Ventral view</term>
<term>Ventrally</term>
<term>Ventromedial</term>
<term>Vertebra</term>
<term>Vertebrate</term>
<term>Vertebrate paleontology</term>
<term>Ypresian</term>
<term>Zoological</term>
<term>Zoological journal</term>
</keywords>
<keywords scheme="Teeft" xml:lang="en">
<term>Abdoun</term>
<term>Abhandlungen</term>
<term>Alveolus</term>
<term>American museum novitates</term>
<term>Anatosuchus</term>
<term>Anterior</term>
<term>Anterior contour</term>
<term>Anterior edge</term>
<term>Anterior margin</term>
<term>Anterior process</term>
<term>Anterior view</term>
<term>Anteriorly</term>
<term>Anteriormost</term>
<term>Anterodorsal</term>
<term>Anterolateral</term>
<term>Anterolateral postorbital process</term>
<term>Anteromedial</term>
<term>Anteromedial margin</term>
<term>Anteroposterior</term>
<term>Anteroposteriorly</term>
<term>Anteroventral</term>
<term>Anteroventrally</term>
<term>Antorbital</term>
<term>Araripesuchus</term>
<term>Archosauria</term>
<term>Arech</term>
<term>Articular</term>
<term>Articular glenoid cavity</term>
<term>Articular surface</term>
<term>Attened</term>
<term>Basal</term>
<term>Basioccipital</term>
<term>Basioccipital tubera</term>
<term>Basisphenoid</term>
<term>Basisphenoid rostrum</term>
<term>Benton</term>
<term>Benton clark</term>
<term>Bequaerti</term>
<term>Bernissartia</term>
<term>Brinkman</term>
<term>Brochu</term>
<term>Broin</term>
<term>Buckley brochu</term>
<term>Buffetaut</term>
<term>Buscalioni</term>
<term>Calsoyasuchus</term>
<term>Calvo</term>
<term>Caudal</term>
<term>Centrum</term>
<term>Cervical</term>
<term>Cervicals</term>
<term>Chenanisuchus</term>
<term>Chenanisuchus lateroculi</term>
<term>Chimaerasuchus</term>
<term>Choanae</term>
<term>Clade</term>
<term>Comahuesuchus</term>
<term>Comptes</term>
<term>Comptes rendus</term>
<term>Concavity</term>
<term>Condyle</term>
<term>Consensus tree</term>
<term>Coracoid</term>
<term>Coronoid</term>
<term>Cranial</term>
<term>Cretaceous</term>
<term>Crocodile</term>
<term>Crocodilia</term>
<term>Crocodilian</term>
<term>Crocodiliens</term>
<term>Crocodyliform</term>
<term>Crocodyliformes</term>
<term>Crocodyliforms</term>
<term>Crocodylomorph</term>
<term>Crocodylomorpha</term>
<term>Crocodylus</term>
<term>Dentary</term>
<term>Dentary teeth</term>
<term>Diapophysis</term>
<term>Dibothrosuchus</term>
<term>Dibothrosuchus protosuchus orthosuchus gobiosuchus notosuchus sebecus uruguaysuchus</term>
<term>Dollo</term>
<term>Dorsal</term>
<term>Dorsal armour</term>
<term>Dorsal margin</term>
<term>Dorsal osteoderms</term>
<term>Dorsal part</term>
<term>Dorsal surface</term>
<term>Dorsal vertebrae</term>
<term>Dorsal view</term>
<term>Dorsally</term>
<term>Dorsals</term>
<term>Dorsoventrally</term>
<term>Dyrosaurid</term>
<term>Dyrosauridae</term>
<term>Dyrosaurids</term>
<term>Dyrosaurus</term>
<term>Dyrosaurus maghribensis</term>
<term>Dyrosaurus phosphaticus</term>
<term>Ectopterygoid</term>
<term>Elosuchus</term>
<term>Eocene</term>
<term>Eustachian</term>
<term>Eutretauranosuchus</term>
<term>Exoccipital</term>
<term>External mandibular fenestra</term>
<term>External nares</term>
<term>External naris</term>
<term>Femur</term>
<term>Fenestra</term>
<term>Foramen</term>
<term>Foramen magnum</term>
<term>Fossa</term>
<term>Gado</term>
<term>Gasparini</term>
<term>Gavialis</term>
<term>Gavialis gangeticus</term>
<term>Glenoid</term>
<term>Glenoid fossa</term>
<term>Gobiosuchus</term>
<term>Gomani</term>
<term>Goniopholis</term>
<term>Goniopholis simus</term>
<term>Halstead</term>
<term>Halstead middleton</term>
<term>Holotype</term>
<term>Hsisosuchus</term>
<term>Humerus</term>
<term>Hyleochampsa</term>
<term>Hypapophyses</term>
<term>Hypapophysis</term>
<term>Hyposaurus</term>
<term>Ianwilsoni</term>
<term>Ilium</term>
<term>Imperator</term>
<term>Infratemporal</term>
<term>Infratemporal fenestra</term>
<term>Infratemporal fenestrae</term>
<term>Interfenestral</term>
<term>Internal nares</term>
<term>Ischium</term>
<term>Jouve</term>
<term>Jouve schwarz</term>
<term>Jugal</term>
<term>Jurassic</term>
<term>Lacrimal</term>
<term>Lamina</term>
<term>Larsson</term>
<term>Larsson gado</term>
<term>Late cretaceous</term>
<term>Lateral</term>
<term>Lateral eustachian foramen</term>
<term>Lateral margin</term>
<term>Lateral margins</term>
<term>Lateral surface</term>
<term>Lateral view</term>
<term>Laterally</term>
<term>Lateroculi</term>
<term>Laterosphenoid</term>
<term>Libycosuchus</term>
<term>Linnean</term>
<term>Linnean society</term>
<term>Longirostrine</term>
<term>Longirostrine forms</term>
<term>Longirostrine morphology</term>
<term>Longitudinal rows</term>
<term>Lower cretaceous</term>
<term>Lower eocene</term>
<term>Lpup lppm</term>
<term>Maghribensis</term>
<term>Malawisuchus</term>
<term>Mandible</term>
<term>Mandibular</term>
<term>Mandibular symphysis</term>
<term>Margin</term>
<term>Maroc</term>
<term>Maxilla</term>
<term>Maxillary</term>
<term>Maxillary teeth</term>
<term>Maxillary tooth</term>
<term>Medial</term>
<term>Medial eustachian foramen</term>
<term>Medial view</term>
<term>Medially</term>
<term>Mera</term>
<term>Mnhn</term>
<term>Monophyly</term>
<term>Mook</term>
<term>Naris</term>
<term>Nasal</term>
<term>Natural history</term>
<term>Neosuchians</term>
<term>Neural</term>
<term>Neural arch</term>
<term>Neural arches</term>
<term>Neural spine</term>
<term>Neural spines</term>
<term>Notosuchia</term>
<term>Notosuchus</term>
<term>Observ</term>
<term>Occipital</term>
<term>Occipital condyle</term>
<term>Occipital face</term>
<term>Occipital tuberosities</term>
<term>Occipital tuberosity</term>
<term>Occipital view</term>
<term>Odontoid process</term>
<term>Orientated</term>
<term>Ortega</term>
<term>Orthosuchus</term>
<term>Osteoderm</term>
<term>Osteoderms</term>
<term>Other crocodyliformes</term>
<term>Other crocodyliforms</term>
<term>Other dyrosaurids</term>
<term>Other longirostrine forms</term>
<term>Otoccipital</term>
<term>Oulad</term>
<term>Oulad abdoun basin</term>
<term>Palaeontology</term>
<term>Palatal</term>
<term>Palatine</term>
<term>Palatine fenestra</term>
<term>Paleontology</term>
<term>Parietal</term>
<term>Paroccipital</term>
<term>Paroccipital process</term>
<term>Patagonicus lomasuchus theriosuchus goniopholis bernissartia hyleochampsa terminonaris pholidosaurus sarcosuchus elosuchus pelagosaurus vectisuchus steneosaurus libycosuchus sunosuchus metriorhynchus dyrosaurus gavialis crocodylus sphenosuchus stolokrosuchus hsisosuchus shamosuchus sichuanosuchus shantungosuchus simosuchus anatosuchus calsoyasuchus rugosuchus alligator kayentasuchus comahuesuchus malawisuchus peirosaurus bretesuchus eutretauranosuchus chenanisuchus sphagesaurus chimaerasuchus</term>
<term>Pelagosaurus</term>
<term>Pers</term>
<term>Pholidosaurus</term>
<term>Phosphaticus</term>
<term>Phylogenetic</term>
<term>Phylogenetic analyses</term>
<term>Phylogenetic analysis</term>
<term>Phylogenetic implications</term>
<term>Phylogenetic position</term>
<term>Phylogenetic relationships</term>
<term>Postcranial</term>
<term>Posterior</term>
<term>Posterior edge</term>
<term>Posterior margin</term>
<term>Posterior margins</term>
<term>Posterior process</term>
<term>Posterior wall</term>
<term>Posteriorly</term>
<term>Posterodorsal</term>
<term>Posterodorsal margin</term>
<term>Posterodorsally</term>
<term>Posterolateral</term>
<term>Posteromedial</term>
<term>Posteromedial process</term>
<term>Posteroventral</term>
<term>Posteroventrally</term>
<term>Postorbital</term>
<term>Postorbital process</term>
<term>Prefrontal</term>
<term>Prefrontal pillar</term>
<term>Prefrontal pillars</term>
<term>Premaxilla</term>
<term>Premaxillary</term>
<term>Present work</term>
<term>Previous investigations</term>
<term>Protosuchus</term>
<term>Proximal head</term>
<term>Pterygoid</term>
<term>Pterygoids</term>
<term>Pubis</term>
<term>Quadrate</term>
<term>Quadrate condyle</term>
<term>Quadratojugal</term>
<term>Rendus</term>
<term>Reptile</term>
<term>Retroarticular</term>
<term>Retroarticular process</term>
<term>Rhabdognathus</term>
<term>Robust</term>
<term>Rogersii</term>
<term>Rostrum</term>
<term>Sacral</term>
<term>Same level</term>
<term>Sanz</term>
<term>Sarcosuchus</term>
<term>Sarcosuchus imperator</term>
<term>Scapula</term>
<term>Schwarz</term>
<term>Sciences naturelles</term>
<term>Sebecus</term>
<term>Sereno</term>
<term>Shantungosuchus</term>
<term>Sichuanosuchus</term>
<term>Simosuchus</term>
<term>Skull</term>
<term>Skull length</term>
<term>Skull roof</term>
<term>Snout</term>
<term>Sphagesaurus</term>
<term>Sphenosuchus</term>
<term>Squamosal</term>
<term>Steneosaurus</term>
<term>Stolokrosuchus</term>
<term>Storrs</term>
<term>Suborbital</term>
<term>Suborbital fenestra</term>
<term>Sunosuchus</term>
<term>Sunosuchus junggarensis</term>
<term>Supraoccipital</term>
<term>Supratemporal</term>
<term>Supratemporal fenestra</term>
<term>Supratemporal fenestrae</term>
<term>Surangular</term>
<term>Suture</term>
<term>Sutured</term>
<term>Symphysis</term>
<term>Taquet</term>
<term>Taxon</term>
<term>Temporal canal</term>
<term>Terminonaris</term>
<term>Thalattosuchia</term>
<term>Thalattosuchians</term>
<term>Theriosuchus</term>
<term>Transverse process</term>
<term>Transverse section</term>
<term>Troxell</term>
<term>Tubera</term>
<term>Tuberosity</term>
<term>Turner calvo</term>
<term>Tykoski</term>
<term>Unpublished dphil thesis</term>
<term>Upper cretaceous</term>
<term>Upper triassic</term>
<term>Uruguaysuchus</term>
<term>Vectisuchus</term>
<term>Vectisuchus leptognathus</term>
<term>Ventral</term>
<term>Ventral edge</term>
<term>Ventral margin</term>
<term>Ventral part</term>
<term>Ventral portion</term>
<term>Ventral view</term>
<term>Ventrally</term>
<term>Ventromedial</term>
<term>Vertebra</term>
<term>Vertebrate</term>
<term>Vertebrate paleontology</term>
<term>Ypresian</term>
<term>Zoological</term>
<term>Zoological journal</term>
</keywords>
<keywords scheme="Wicri" type="geographic" xml:lang="fr">
<term>Maroc</term>
</keywords>
<keywords scheme="Wicri" type="topic" xml:lang="fr">
<term>Reptile</term>
</keywords>
</textClass>
</profileDesc>
</teiHeader>
<front>
<div type="abstract">A new dyrosaurid is described from the Ypresian of the phosphatic deposits of the Oulad Abdoun Basin of Morocco. It is based on numerous cranial and postcranial remains, allowing an almost complete reconstruction. This new Dyrosaurus species, Dyrosaurus maghribensis sp. nov., is currently only known from Morocco. It differs from D. phosphaticus, present in contemporaneous levels of Algeria and Tunisia, by several autapomorpies, including a smooth dorsal margin of the parietal and widely opened choanae. A phylogenetic analysis, using 47 taxa and 234 morphological characters, shows the dyrosaurids as the sister taxon of pholidosaurids, which include Elosuchus, Sarcosuchus, Terminonaris and Pholidosaurus, and the thalattosuchians. Goniopholididae is a non‐monophyletic group; however, if dyrosaurids are not included in the analysis, the result differs and the goniopholidids form a distinct clade. If Thalattosuchia is excluded, both Goniopholididae and Pholidosauridae become paraphyletic assemblages. Thus, phylogenetic problems remain with respect to longirostrine clade, and more attention should be paid to resolving their evolutionary relationships amongst the crocodyliforms. © 2006 The Linnean Society of London, Zoological Journal of the Linnean Society, 2006, 148, 603–656.</div>
</front>
</TEI>
<affiliations>
<list>
<country>
<li>France</li>
<li>Maroc</li>
</country>
<region>
<li>Rabat-Salé-Kénitra</li>
<li>Île-de-France</li>
</region>
<settlement>
<li>Paris</li>
<li>Rabat</li>
</settlement>
</list>
<tree>
<country name="France">
<region name="Île-de-France">
<name sortKey="Jouve, Stephane" sort="Jouve, Stephane" uniqKey="Jouve S" first="Stéphane" last="Jouve">Stéphane Jouve</name>
</region>
<name sortKey="Jouve, Stephane" sort="Jouve, Stephane" uniqKey="Jouve S" first="Stéphane" last="Jouve">Stéphane Jouve</name>
</country>
<country name="Maroc">
<region name="Rabat-Salé-Kénitra">
<name sortKey="Iarochene, Mohamed" sort="Iarochene, Mohamed" uniqKey="Iarochene M" first="Mohamed" last="Iarochène">Mohamed Iarochène</name>
</region>
<name sortKey="Amaghzaz, Mbarek" sort="Amaghzaz, Mbarek" uniqKey="Amaghzaz M" first="Mbarek" last="Amaghzaz">Mbarek Amaghzaz</name>
<name sortKey="Bouya, Baadi" sort="Bouya, Baadi" uniqKey="Bouya B" first="Baâdi" last="Bouya">Baâdi Bouya</name>
</country>
</tree>
</affiliations>
</record>

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